Polarity and connectivity of DAL neurons. New protein synthesis for LTM occurs in the two dorsal–anterior–lateral (DAL) neurons (magenta) with axonal terminals (light blue) linking to pioneer a/b neurons (green) at the K5 dendritic field (white) in the calyx. The DAL neuron may receive inputs from dorsal frontal protocerebrum where it has numerous dendrites (yellow). The brain is counterstained with anti-disc large immunostaining (grey).